اثنی عشری، م. و زکایی خسروشاهی، م. ۱۳۸۷. پلیآمینها و علوم باغبانی. انتشارات دانشگاه بوعلی سینا، ۱۹۳ ص.
حاج سیدحاجی، م.ر. و رضایی قلعه، ه. 1394. بررسی تأثیر مقادیر مختلف ورمیکمپوست و محلولپاشی اسیدهای آمینه و اوره بر عملکرد کمّی و کیفی بابونه آلمانی (Matricaria chamomilla L.). تحقیقات گیاهان دارویی و معطر ایران، 31(6): 1058-1070.
خسروشاهی، م. 1386. اثر پوترسین بر عمر پس از برداشت میوههای توتفرنگی، زردآلو، هلو و گیلاس. مجله علوم و فنون کشاورزی، 12(45): 228-219.
خوشخوی، م.، شیبانی، ب.، روحانی، ا. و تفضلی، ع. ۱۳۸۹. اصول باغبانی. انتشارات دانشگاه شیراز، چاپ نوزدهم، 596 ص.
داورپناه، س.، عسگریسرچشمه، م.، بابالار، م و حسینی، م. 1392. بررسی ضرورت تغذیه برگی برخی از عناصر در خاکهای آهکی. اولین همایش ملی الکترونیکی مباحث نوین در علوم باغبانی، جهرم.
مجد، ا.، صفاری، پ.، جنوبی، پ و مهرابیان، ص. 1390. بررسی مراحل تکوینی بخشهای رویشی و زایشی انگور شاهانی. فصلنامه علمی پژوهشی زیستشناسی تکوینی، 3(10): 61-51.
ملکوتی، م و همایی، م. ۱۳۸۳. حاصلخیزی خاکهای مناطق خشک و نیمه خشک. انتشارات دانشگاه تربیت مدرس. ۸۱ ص.
نیکبخت، ع.، کافی، م.، بابالار، م.، اعتمادی، ن.، ابراهیمزاده، ح. و پینگشی، ی. ۱۳۸۹. اثر هیومیک اسید بر جذب کلسیم و رفتار فیزیولوژیکی پس از برداشت گل ژربرا، مجله علوم و فنون باغبانی ایران، ۸(4): 268-237.
Aebi, H., 1984. Catalase in vitro. In Methods in enzymology (Vol. 105, pp. 121-126). Academic press.
Akinci, S., Bueyuekkeskin, T., Eroglu, A. and Erdogan, B.E. 2009. The effect of humic acid on nutrient composition in broad bean (Vicia faba L.) roots. Notulae Scientia Biologicae, 1(1): 81-87.
Alcázar, R., Bueno, M. and Tiburcio, A.F. 2020. Polyamines: Small amines with large effects on plant abiotic stress tolerance. Cells, 9(11): 2373.
Ali, B., Wang, X., Saleem, M.H., Sumaira, Hafeez, A., Afridi, M.S., Khan, S., Ullah, I., Amaral Júnior, A.T.D., Alatawi, A. and Ali, S. 2022. PGPR-mediated salt tolerance in maize by modulating plant physiology, antioxidant defense, compatible solutes accumulation and bio-surfactant producing genes. Plants, 11(3): 345.
Bayat, H., Shafie, F., Aminifard, M.H. and Daghighi, S. 2021. Comparative effects of humic and fulvic acids as biostimulants on growth, antioxidant activity and nutrient content of yarrow (Achillea millefolium L.). Scientia Horticulturae, 279: 109912.
Braziene, Z., Paltanavicius, V. and Avizienytė, D. 2021. The influence of fulvic acid on spring cereals and sugar beets seed germination and plant productivity. Environmental Research, 195:110824.
Bronick, C.J. and Lal, R. 2005. Soil structure and management: a review. Geoderma, 124(1-2): 3-22.
Chen, J., Fang, J., Guo, Z. and Lu, S. 2018. Polyamines and antioxidant defense system are associated with cold tolerance in centipedegrass. Frontiers of Agricultural Science and Engineering, 5(1): 129-138.
Choudhary, S., Wani, K.I., Naeem, M., Khan, M.M.A. and Aftab, T. 2023. Cellular responses, osmotic adjustments, and role of osmolytes in providing salt stress resilience in higher plants: polyamines and nitric oxide crosstalk. Journal of Plant Growth Regulation, 42(2): 539-553.
Cordeiro, F.C., Santa-Catarina, C., Silveira, V. and De Souza, S.R., 2011. Humic acid effect on catalase activity and the generation of reactive oxygen species in corn (Zea mays). Bioscience, Biotechnology, and Biochemistry, 75(1): 70-74.
Desoky, E.S.M., Merwad, A.R.M., Abo El-Maati, M.F., Mansour, E., Arnaout, S.M., Awad, M.F., Ramadan, M.F. and Ibrahim, S.A. 2021. Physiological and biochemical mechanisms of exogenously applied selenium for alleviating destructive impacts induced by salinity stress in bread wheat. Agronomy, 11(5): 926.
Elmongy, M.S., Zhou, H., Cao, Y., Liu, B. and Xia, Y. 2018. The effect of humic acid on endogenous hormone levels and antioxidant enzyme activity during in vitro rooting of evergreen azalea. Scientia Horticulturae, 227: 234-243.
ElSayed, A.I., Mohamed, A.H., Rafudeen, M.S., Omar, A.A., Awad, M.F. and Mansour, E. 2022. Polyamines mitigate the destructive impacts of salinity stress by enhancing photosynthetic capacity, antioxidant defense system and upregulation of calvin cycle-related genes in rapeseed (Brassica napus L.). Saudi Journal of Biological Sciences, 29(5): 3675-3686.
Esfandiari Ghalati, R., Shamili, M. and Homaei, A. 2020. Effect of putrescine on biochemical and physiological characteristics of guava (
Psidium guajava L.) seedlings under salt stress.
Scientia Horticulturae, 261: 108961.
Eshwar, M., Srilatha, M., Rekha, K.B., Sharma, S.H.K. and Eshwar, C.M. 2017. Effect of humic substances (humic, fulvic acid) and chemical fertilizers on nutrient uptake, dry matter production of aerobic rice (Oryza sativa L.). Journal of Pharmacognosy and Phytochemistry, 6(5): 1063-1066.
Evelin, H., Devi, T.S., Gupta, S. and Kapoor, R. 2019. Mitigation of salinity stress in plants by arbuscular mycorrhizal symbiosis: current understanding and new challenges. Frontiers in Plant Science, 10: 470.
Fathy, M.A., Gabr, M.A. and El Shall, S.A. 2010. Effect of humic acid treatments on'Canino'apricot growth, yield and fruit quality. New York Science Journal, 3(12): 109-115.
Gai, Z., Liu, L., Zhang, J., Liu, J. and Cai, L. 2020. Effects of exogenous α-oxoglutarate on proline accumulation, ammonium assimilation and photosynthesis of soybean seedling (Glycine max (L.) Merr.) exposed to cold stress. Scientific Reports, 10(1): 17017.
Ghosh, U.K., Islam, M.N., Siddiqui, M.N. and Khan, M.A.R., 2021. Understanding the roles of osmolytes for acclimatizing plants to changing environment: a review of potential mechanism. Plant Signaling and Behavior, 16(8):1913306.
Giovanardi, D., Dallai, D., Dondini, L., Mantovani, V. and Stefani, E. 2016. Elicitation of resistance to bacterial canker of stone fruits by humic and fulvic acids (glucohumates): a cDNA-AFLP-dHPLC approach. Scientia Horticulturae, 212: 183-192.
Gong, X., Liu, Y., Huang, D., Zeng, G., Liu, S., Tang, H., Zhou, L., Hu, X., Zhou, Y. and Tan, X. 2016. Effects of exogenous calcium and spermidine on cadmium stress moderation and metal accumulation in Boehmeria nivea (L.) Gaudich. Environmental Science and Pollution Research, 23: 8699-8708.
Habba, E.E., Abdel Aziz, N.G., Sarhan, A.M.Z., Arafa, A.M.S. and Youssefi, N.M. 2016. Effect of putrescine and growing media on vegetative growth and chemical constituents of Populus euramericana plants. Journal of Innovations in Pharmaceutical and Biological Sciences, 3(1): 61-73.
Hasanuzzaman, M., Parvin, K., Bardhan, K., Nahar, K., Anee, T.I., Masud, A.A.C. and Fotopoulos, V., 2021. Biostimulants for the regulation of reactive oxygen species metabolism in plants under abiotic stress. Cells, 10(10): 2537.
Hussain, A., Nazir, F. and Fariduddin, Q. 2019. 24-epibrassinolide and spermidine alleviate Mn stress via the modulation of root morphology, stomatal behavior, photosynthetic attributes and antioxidant defense in Brassica juncea. Physiology and Molecular Biology of Plants, 25(4): 905-919.
Hwang, I., Sheen, J. and Müller, B., 2012. Cytokinin signaling networks. Annual Review of Plant Biology, 63(1): 353-380.
Irigoyen, J.J., Einerich, D.W. and Sánchez‐Díaz, M. 1992. Water stress induced changes in concentrations of proline and total soluble sugars in nodulated alfalfa (Medicago sativa) plants. Physiologia Plantarum, 84(1): 55-60.
Islam, M.A., Morton, D.W., Johnson, B.B. and Angove, M.J. 2020. Adsorption of humic and fulvic acids onto a range of adsorbents in aqueous systems, and their effect on the adsorption of other species: A review. Separation and Purification Technology, 247:116949.
Jankovska-Bortkevič, E., Jurkonienė, S., Gavelienė, V., Šveikauskas, V., Mockevičiūtė, R., Vaseva, I., Todorova, D., Žižytė-Eidetienė, M., Šneideris, D. and Prakas, P. 2023. Dynamics of Polyamines, proline, and ethylene metabolism under increasing cold in winter oilseed rape. International Journal of Molecular Sciences, 24(14): 11402.
Jia, X.M., Zhu, Y.F., Hu, Y., Zhang, R., Cheng, L., Zhu, Z.L., Zhao, T., Zhang, X. and Wang, Y.X. 2019. Integrated physiologic, proteomic, and metabolomic analyses of Malus halliana adaptation to saline–alkali stress. Horticulture Research, 6: 91.
Jiao, X., Li, Y., Zhang, X., Liu, C., Liang, W., Li, C., Ma, F. and Li, C. 2019. Exogenous dopamine application promotes alkali tolerance of apple seedlings. Plants, 8(12): 580.
Juurakko, C.L. and Walker, V.K. 2021. Cold acclimation and prospects for cold-resilient crops. Plant Stress, 2: 100028.
Kang, H.M. and Saltveit, M.E. 2002. Chilling tolerance of maize, cucumber and rice seedling leaves and roots are differentially affected by salicylic acid. Physiologia Plantarum, 115(4): 571-576.
Karimi, R. and Salimi, F. 2021. Iron-chlorosis tolerance screening of 12 commercial grapevines (Vitis vinifera L.) cultivars based on phytochemical indices. Scientia Horticulturae, 283: 110111.
Karimi, E., Tadayyon, A. and Tadayyon, M.R. 2016. The effect of humic acid on some yield characteristics and leaf proline content of safflower under different irrigation regimes. Journal of Crops Improvement, 18(3): 609-623.
Kaur-Sawhney, R., Tiburcio, A.F., Altabella, T. and Galston, A.W. 2003. Polyamines in plants: an overview. Journal of Cell and Molecular Biology, 2:1-12.
Kocher, G.S. and Nikhanj, P. 2019. Development of red and white wines from locally adapted grape cultivars using indigenous yeast. In Fermented Beverages (pp. 147-170). Woodhead Publishing.
Kusano, T., Berberich, T., Tateda, C. and Takahashi, Y. 2008. Polyamines: essential factors for growth and survival. Planta, 228: 367-381.
Lechowska, K., Wojtyla, Ł., Quinet, M., Kubala, S., Lutts, S. and Garnczarska, M. 2021. Endogenous polyamines and ethylene biosynthesis in relation to germination of osmoprimed Brassica napus seeds under salt stress. International Journal of Molecular Sciences, 23(1): 349.
Lichtenthaler, H.K. 1987. Chlorophylls and carotenoids: pigments of photosynthetic biomembranes. In Methods in enzymology (Vol. 148, pp. 350-382). Academic Press.
Liu, J.H., Wang, W., Wu, H., Gong, X. and Moriguchi, T. 2015. Polyamines function in stress tolerance: from synthesis to regulation. Frontiers in plant science, 6: 827.
Liu, X., Yang, J., Tao, J. and Yao, R. 2022. Integrated application of inorganic fertilizer with fulvic acid for improving soil nutrient supply and nutrient use efficiency of winter wheat in a salt-affected soil. Applied Soil Ecology, 170: 104255.
Lu, X., Ma, L., Zhang, C., Yan, H., Bao, J., Gong, M., Wang, W., Li, S., Ma, S. and Chen, B. 2022. Grapevine (Vitis vinifera) responses to salt stress and alkali stress: transcriptional and metabolic profiling. BMC Plant Biology, 22(1): 528.
Movahed, N., Eshghi, S., Tafazoli, E. and Jamali, B. 2010. Effects of polyamines on vegetative characteristics, growth, flowering and yield of strawberry ('PAROS'AND'SELVA'). In XXVIII International Horticultural Congress on Science and Horticulture for People (IHC2010): International Symposium, 926: 287-293.
Mustafavi, S.H., Naghdi Badi, H., Sękara, A., Mehrafarin, A., Janda, T., Ghorbanpour, M. and Rafiee, H. 2018. Polyamines and their possible mechanisms involved in plant physiological processes and elicitation of secondary metabolites. Acta Physiologiae Plantarum, 40:1-19.
Naeem, M., Shahzad, K., Saqib, S., Shahzad, A., Nasrullah, Younas, M. and Afridi, M.I. 2022. The Solanum melongena COP1LIKE manipulates fruit ripening and flowering time in tomato (Solanum lycopersicum). Plant Growth Regulation, 96: 369-382.
Nahed, G.A.A., Lobna, S.T. and Soad, M.I. 2009. Some studies on the effect of putrescine, ascorbic acid and thiamine on growth, flowering and some chemical constituents of gladiolus plants at Nubaria. Ozean Journal of Applied Sciences, 2(2):169-179.
Najm, A.A., Hadi, M.R.H.S., Fazeli, F., Darzi, M.T. and Rahi, A. 2012. Effect of integrated management of nitrogen fertilizer and cattle manure on the leaf chlorophyll, yield, and tuber glycoalkaloids of Agria potato. Communications in Soil Science and Plant Analysis, 43(6): 912-923.
Nakano, Y. and Asada, K. 1981. Hydrogen peroxide is scavenged by ascorbate-specific peroxidase in spinach chloroplasts. Plant and Cell Physiology, 22(5): 867-880.
Nardi, S., Pizzeghello, D., Muscolo, A. and Vianello, A. 2002. Physiological effects of humic substances on higher plants. Soil Biology and Biochemistry, 34(11): 1527-1536.
Neily, M.H., Matsukura, C., Maucourt, M., Bernillon, S., Deborde, C., Moing, A., Yin, Y.G., Saito, T., Mori, K., Asamizu, E. and Rolin, D. 2011. Enhanced polyamine accumulation alters carotenoid metabolism at the transcriptional level in tomato fruit over-expressing spermidine synthase. Journal of Plant Physiology, 168(3): 242-252.
Ozturk, M., Turkyilmaz Unal, B., García‐Caparrós, P., Khursheed, A., Gul, A. and Hasanuzzaman, M. 2021. Osmoregulation and its actions during the drought stress in plants. Physiologia Plantarum, 172(2): 1321-1335.
Paquin, R. and Lechasseur, P. 1979. Observations ssue une methode de dosage de la proline libre dans les extraits de plantes. Canadian Journal of Botany, 75:1851-1854.
Paul, S., Banerjee, A. and Roychoudhury, A. 2018. Role of polyamines in mediating antioxidant defense and epigenetic regulation in plants exposed to heavy metal toxicity. In Plants under metal and metalloid stress: Responses, tolerance and remediation. Singapore: Springer Singapore, pp. 229-247.
Ramazan, S., Nazir, I., Yousuf, W. and John, R. 2022. Environmental stress tolerance in maize (Zea mays): role of polyamine metabolism. Functional Plant Biology, 50(2): 85-96.
Roosta, H.R., Manzari Tavakkoli, M. and Hamidpour, M. 2016. Comparison of different soilless media for growing gerbera under alkalinity stress condition. Journal of Plant Nutrition, 39(8): 1063-1073.
Şahin, Ö., Dumanoğlu, H., Sarıkamış, G., Javadisaber, J., Ergül, A. and Aydemir, B.Ç. 2022. Tolerance of Pyrus spp. and Cydonia oblonga as pear rootstocks to iron chlorosis determined by in vitro growth, antioxidant and molecular responses. Scientia Horticulturae, 296: 110911.
Saleem, A., Zulfiqar, A., Saleem, M.Z., Ali, B., Saleem, M.H., Ali, S., Tufekci, E.D., Tufekci, A.R., Rahimi, M. and Mostafa, R.M. 2023. Alkaline and acidic soil constraints on iron accumulation by Rice cultivars in relation to several physio-biochemical parameters. BMC Plant Biology, 23(1): 397.
Sattar, A., Wang, X., Abbas, T., Sher, A., Ijaz, M., Ul-Allah, S., Irfan, M., Butt, M., Wahid, M.A., Cheema, M. and Fiaz, S. 2021. Combined application of zinc and silicon alleviates terminal drought stress in wheat by triggering morpho-physiological and antioxidants defense mechanisms. PloS one, 16(10): 0256984.
Shahsavandi, F. and Eshghi, S. 2021. Effects of bicarbonate and Fe sources on vegetative growth and physiological traits of four grapevine cultivars. Communications in Soil Science and Plant Analysis, 52(20): 2401-2413.
Sharif, M., Khattak, R.A. and Sarir, M.S. 2002. Effect of different levels of lignitic coal derived humic acid on growth of maize plants. Communications in Soil Science and Plant Analysis, 33(19-20): 3567-3580.
Suh, H.Y., Yoo, K.S. and Suh, S.G. 2014. Effect of foliar application of fulvic acid on plant growth and fruit quality of tomato (Lycopersicon esculentum L.). Horticulture, Environment, and Biotechnology, 55: 455-461.
Sun, M., Wang, T., Fan, L., Wang, H., Pan, H., Cui, X., Lou, Y. and Zhuge, Y. 2020. Foliar applications of spermidine improve foxtail millet seedling characteristics under salt stress. Biologia Plantarum, 64(1).
Szabados, L. and Savouré, A. 2010. Proline: a multifunctional amino acid. Trends in Plant Science, 15(2): 89-97.
Talaat, I.M., Bekheta, M.A. and Mahgoub, M.H. 2005. Physiological response of periwinkle plants (Catharanthus roseus L.) to tryptophan and putrescine. International Journal of Agricultural and Biological Engineering, 7(2): 210-213.
Thialla, L.A., Damiana, C.M., Arthur, A.S.O., Reinaldo, A.P., Walter, S.E.J. and Djair, A.M. 2017. Gibberellin and polyamines in plant growth, development, and postharvest senescence of ornamental plants. Amazonian Journal of Plant Research, 1(10): 1-13.
Tripathi, D.K., Bhat, J.A., Ahmad, P. and Allakhverdiev, S.I., 2023. Polyamines and nitric oxide crosstalk in plant development and abiotic stress tolerance. Functional Plant Biology, 50(2): 1-4.
Tyagi, A., Ali, S., Ramakrishna, G., Singh, A., Park, S., Mahmoudi, H. and Bae, H. 2023. Revisiting the role of polyamines in plant growth and abiotic stress resilience: mechanisms, crosstalk, and future perspectives. Journal of Plant Growth Regulation, 42(8): 5074-5098.
Unlu, H. O., Unlu, H., Karakurt, Y. and Padem, H. 2011. Changes in fruit yield and quality in response to foliar and soil humic acid application in cucumber. Scientific Research and Essays, 6(13): 2800-2803.
Upadhyaya, A., Sankhla, D., Davis, T.D., Sankhla, N. and Smith, B.N. 1985. Effect of paclobutrazol on the activities of some enzymes of activated oxygen metabolism and lipid peroxidation in senescing soybean leaves. Journal of Plant Physiology, 121(5): 453-461.
Valdez-Aguilar, L.A. and Reed, D.W. 2008. Influence of potassium substitution by rubidium and sodium on growth, ion accumulation, and ion partitioning in bean under high alkalinity. Journal of Plant Nutrition, 31(5): 867-883.
Valero, D., Martı́nez-Romero, D. and Serrano, M. 2002. The role of polyamines in the improvement of the shelf life of fruit. Trends in Food Science and Technology, 13(6-7): 228-234.
Valipour, M., Baninasab, B., Khoshgoftarmanesh, A.H. and Gholami, M. 2020. Oxidative stress and antioxidant responses to direct and bicarbonate-induced iron deficiency in two quince rootstocks. Scientia Horticulturae, 261: 108933.
Verbruggen, N. and Hermans, C. 2008. Proline accumulation in plants: A review. Amino Acids, 35: 753-759.
Wallace, H.M. 2009. The polyamines: past, present and future. Essays in Biochemistry, 46: 1-10.
Wu, J., Shu, S., Li, C., Sun, J. and Guo, S. 2018. Spermidine-mediated hydrogen peroxide signaling enhances the antioxidant capacity of salt-stressed cucumber roots. Plant Physiology and Biochemistry, 128: 152-162.
Qiang-Sheng, W.U., Ying-Ning, Z.O.U., Min, L.I.U. and Cheng, K. 2012. Effects of exogenous putrescine on mycorrhiza, root system architecture, and physiological traits of Glomus mosseae-colonized trifoliate orange seedlings. Notulae Botanicae Horti Agrobotanici Cluj-Napoca, 40(2): 80-85.
Xuan, T.D., Huong, C.T., Quan, N.V., Anh, L.H., Khanh, T.D. and Rayee, R. 2022. Improvement of salinity tolerance in rice seedlings by exogenous magnesium sulfate application. Soil Systems, 6(3): 69.
Zhang, M., Li, X., Wang, X., Feng, J. and Zhu, S. 2023. Potassium fulvic acid alleviates salt stress of citrus by regulating rhizosphere microbial community, osmotic substances and enzyme activities. Frontiers in Plant Science, 14:1161469.
Zheng, X., Wang, L., Zhou, J., Shan, D., Wang, N. and Kong, J. 2021. Melatonin enhances the tolerance to iron deficiency stress through scavenging ROS in apple. Fruit Research, 1(1): 1-8.
Zhu, H., Tian, W., Zhu, X., Tang, X., Wu, L., Hu, X. and Jin, S. 2020. Ectopic expression of GhSAMDC 1 improved plant vegetative growth and early flowering through conversion of spermidine to spermine in tobacco. Scientific Reports, 10(1): 14418.